Abbink, O.A., 1998. Palynological investigations in the Jurassic of the North Sea region. Ph.D Thesis,
University of Utrecht, LPP Contribution Series, 8: 192 p.
Abbink, O., Targarona, J., Brinkhuis, H., Visscher, H., 2001. Late Jurassic to earliest Cretaceous
palaeoclimatic evolution of the southern North Sea. Global and Planetary Change, 30(3-4): 231-256.
Abbink, O.A., van Konijnenburg-van Cittert, J.H.A., van der Zwan, C.J., Visscher, H., 2004. A
sporomorph ecogroup model for the Northwest European Jurassic-Lower Cretaceous II: Application
to an exploration well from the Dutch North Sea. Netherlands Journal of Geosciences, 83(2): 81-91.
Achilles, H., Kaiser, H., Schweitzer, H.J., 1984. Die rato-jurassischen Floren des Iran und Afghanistans.
7. Die Microflora der obertriadischjurassischen Ablagerungen des Alborz-Gebirges (Nord-Iran).
Palaeontographica B, 194 (1-4): 14-95.
Agard, P., Omrani, J., Jolivet, J., Whiterurch, H., Vrielynck, B., Spakman, W., Monie, P., Meyer, B.,
Wortel, R., 2011. Zagros orogeny: a subduction dominated process. Geological Magazine, 148: 692-
725. DOI:10.1017/S001675681100046X.
Aghanabati, S.A., 1998. Jurassic stratigraphy of Iran, vol. 1. Tehran. Geological Survey of Iran, 355 pp.
(in Persian).
Aghanabati, S.A., 2014. Stratigraphic Lexicon of Iran, Volume 4: Jurassic. Geological Survey of Iran,
544 pp.
374 Vaez-Javadi et al.
Aghanabati, S.A., Haghipour, A., 1978. Geological map of Tabas, 1:250000, No. 17, Report 3(4),
Tehran. Geological Survey of Iran (in Persian).
Alavi, M., Vaziri, H., Seyed-Emami, K., Lasemi, Y., 1977. The Triassic and associated rocks of the
Nakhlak and Aghdarband areas in central and northeastern Iran as remnants of the southern Turanian
active continental margin. Geological Society of America Bulletin, 109: 1563-1575.
Alvin, K.L., 1982. Cheirolepidiaceae - biology, structure and paleo-ecology. Review of Palaeobotany
and Palynology, 37: 71-98.
Ameri, H., Dastanpour, M., Khalilizade, H., Zamani, F., 2014. Plant fossil remains from the Bajocian-
Bathonian of Hojedk formation, Babhutk area, Kerman, Iran. Arabian Journal of Geosciences, 7 (6):
2293-2302.
Arjang, B., 1975. Die räto-jurassischen Floren des Iran und Afghanistans. 1. Die Microflora der ratojurassischen
Ablagerungen des Kermaner Beckens (Zentral Iran). Palaeontographica B, 152 (4-6):
85-148.
Ashraf, A.R., 1977. Die räto-jurassischen Floren des Iran und Afghanistans. 3. Die Mikrofloren der
ratischen bis unterkretazischen Ablagerungen Nordafghanistans. Palaeontographica B, 161 (1-4): 1-
97.
Badihagh, MT., Sajjadi, F., Farmani, T., Uhl, D., 2019. Middle Jurassic palaeoenvironment and
palaeobiogeography of the Tabas Block, Central Iran: palynological and palaeobotanical
Investigations. Palaeobiodiversity and Palaeoenvironments, 99: 379-399.
Balme, B.E., 1995. Fossil in situ spores and pollen grains: An annotated catalogue. Review of
Palaeobotany and Palynology, 87: 1-323.
Barbacka, M., 2009. Sphenophyta from the Early Jurassic of the Mecsek Mts., Hungary. Acta
Palaeobotanica, 49: 221-231.
Barbacka, M., Bodor, E., Jarzynka, A., Kustatscher, E., Pacyna, G., Popa, M. E., Scanu, G. G.,
Thévenard, F., Ziaja, J., 2014. European Jurassic floras: statistics and palaeoenvironmental proxies.
Acta Palaeobotanica, 54: 173-195.
Batten, D.J., 1974. Wealden palaeoecology from the distribution of plant fossils. Proceedings of the
Geological Association, 85(4): 433-458.
Beckett, P.H.T., 1956. Coal deposits near Kirman, south Persia. Economic Geology, 51(2): 197-198.
Berberian, F. & Berberian, M., 1981. Tectono-Plutonic Episodes in Iran. Geological Survey of Iran,
Report 52: 566-593.
Berberian, M., King, G.C.P., 1981. Towards a paleogeography and tectonic evolution of Iran. Canadian
Journal of Earth Sciences, 18 (2): 210-265.
Berra, F., Zanchi, A., Angiolini, L., Vachard, D., Vezzoli, G., Zanchetta, S. ... and Kouhpeyma, M.,
2017. The upper Paleozoic Godar-e-Siah Complex of Jandaq: evidence and significance of a North
Palaeotethyan succession in Central Iran. Journal of Asian Earth Sciences, 138: 272-290.
Bharadwaj, D.C. & Kumar, P., 1986. Palynology of Jurassic sediments from Iran: vol. 1, Kerman area.
Biological Memoirs, 12 (2): 146-172.
Bruun Christensen, O., 1995. Mid-Late Jurassic palaeoenvironments in the Northern North
Sea, Norway as characterised by macroflora and -fauna elements. Geobios, 28: 69-76.
Christenhusz, M.J.M., Reveal, J.L., Farjon, A., Gardner, M.F., Mill, R.R., Chase, M.W., 2011. A new
classification and linear sequence of extant gymnosperms. Phytotaxa, 19: 55-70.
Costamagna, L.G., Kustatscher, E., Scanu, G.G., Del Rio M., Pittau, P., van Konijnenburg-van Cittert,
J.H.A., 2018. A palaeoenvironmental reconstruction of the Middle Jurassic of Sardinia (Italy) based
on integrated palaeobotanical, palynological and lithofacies data assessment. Palaeobiodiversity and
Palaeoenvironments, 98 (1): 111-138.
Davoudzadeh, M., Soffel, H., Schmidt, K., 1981. On the rotation of Central-East-Iran microplate. Neues
Jahrbuch Geologie und Paläontologie, 3: 180-192.
Decombeix, A.L., Bomfleur, B., Taylor, E.L., Taylor, T.N., 2014. New insights into the anatomy,
development, and affinities of corystosperm trees from the Triassic of Antarctica. Review of
Palaeobotany and Palynology, 203: 22-34.
Dehbozorgi, A., Sajjadi, F., Hashemi, H., 2013. Middle Jurassic palynomorphs of the Dalichai
Formation, central Alborz Ranges, northeastern Iran, Paleoecological inferences. Science China
Earth Sciences, 56 (12): 2107-2115. DOI: 10.1007/s11430-013-4697-z.
De Jersey, N.J., 1970. Triassic miospores from the Blackstone Formation, Aberdare Conglomerate and
Geopersia 2024, 14(2): 361-377 375
Raceview Formation. Geological Survey of Queensland Publications, 348: 41.
Fotoohi Rad, G.R., Droop, G.T.R., Burgess, R., 2009. Early cretaceous exhumation of high-pressure
metamorphic rocks of the Sistan Suture Zone, eastern Iran. Geological Journal, 44 (1): 104-116.
Francis, J.E., 1983. The dominant conifer of the Jurassic Purbeck Formation, England.
Palaeontology, 26: 277-294.
Götz, A.E., Ruckwied, K., Barbacka, M., 2011. Paleoenvironment of the Late Triassic (Rhaetian) and
Early Jurassic (Hettangian) Mecsek Coal Formation (south Hungary): implication from macro and
microfloral assemblage. Palaeogeography, Palaeoclimatology, Palaeoecology, 91: 75-88.
Harris, T.M., 1964. The Yorkshire Jurassic Flora. Vol. II: Caytoniales, Cycdales & Pteridosperms.
Trustees of the British Museum (Natural History), London, 191 pp.
Harris, T.M., 1983. The stem of Pachypteris papillosa (Thomas and Bose) Harris. Botanical Journal of
the Linnean Society, 86: 149-159.
Hashemi Yazdi, F., Sajjadi, F.,& Hashemi, H., 2014. Palynostratigraphy of the Hojedk Formation in the
Eshkelli section, north of Kerman based on miospores. Journal of Paleontology, 1 (2): 111-127 (in
Persian).
Hill, C.R., 1990. Ultrastructure of in situ fossil Cycad pollen from the English Jurassic, with a
description of the male cone Androstrobus balmei sp. nov. Review of Palaeobotany and Palynology,
65: 165-173.
Khoddam-Alhoseini, S.R., Razavi, B., Jarahi, A., 1989. Primarily report of the exploration of thermal
coals in the North Kouchekali area. Tabas: Ministry of Metals and Mines [unpublished report].
Khorrami, F., Vernant, P., Masson, F., Nilfouroushan, F., Mousavi, Z., Nankali, H., Saadat, S.A.,
Walpersdorf, A., Hosseini, S., Tavakoli, P., Aghamohammadi, A., Alijanzade, M., 2019. An up-todate
crustal deformation map of Iran using integrated campaign-mode and permanent GPS velocities.
Geophysical Journal International, 217: 832-843. https://doi.org/10.1093/gji/ggz045.
Kimyai, A., 1968. Jurassic plant microfossils from the Kerman region. Bulletin of Iranian Petroleum
Institute, 33: 3-23.
Kimyai, A., 1974. Jurassic plant microfossils from Iran. Birbal Sahni Institute of Palaeobotany, Special
Publication, 3: 1-8.
Kimyai, A., 1975. Jurassic palynological assemblages from the Shahrud region, Iran. Geoscience and
Man, 11: 117-121.
Kimyai, A., 1977. Further information on the palynological stratigraphy of the Mesozoic coaly
sediments from Kerman, Iran. Iranian Petroleum Institute, Proceedings of 2nd Geolological
Symposium of Iran, Tehran, 191-217 (in Persian).
Kershaw, P., Wagstaff, B., 2001. The southern conifer family Araucariaceae: history, status, and value
for paleoenvironmental reconstruction. Annual Review of Ecology and Systematics, 32: 397-414.
Kramer, K.U. & Green, P.S., 1990. The Families and Genera of Vascular Plants: Pteridophytes and
Gymnosperms. Springer Berlin, Heidelberg, 404 pp.
Krassilov, V.A., 1977. Contributions to the knowledge of Caytoniales. Review of Palaeobotany and
Palynology, 24: 155-178.
Krassilov, V.A., 2009. Diversity of Mesozoic gnetophytes and the first angiosperms. Paleontology
Journal, 43: 1272-1280.
Kremp, G.O.W., Ames, H.T., Kovar, A.J., 1960a. Catalog of fossil spores and pollen. Volume 9: Triassic
and Jurassic Spores and Pollen. University Park, Pennsylvania.
Kremp, G.O.W. & Ames, H.T., 1961b. Catalog of fossil spores and pollen. Volume 14: Mesozoic and
Tertiary Spores and Pollen. University Park, Pennsylvania.
Kremp, G.O.W. & Ames, H.T., 1962a. Catalog of fossil spores and pollen. Volume 15: Tertiary and
Cretaceous Spores and Pollen. University Park, Pennsylvania.
Kustatscher, E., van Konijnenburg-van Cittert, J.H.A., Roghi, G., 2010. Macrofloras and palynomorphs
as possible proxies for palaeoclimatic and palaeoecological studies: A case study from the Pelsonian
(Middle Triassic) of Kuhwiesenkopf/Monte Pra della Vacca (Olang Dolomites, N-Italy).
Palaeogeography, Palaeoclimatology, Palaeoecology, 290: 71-80.
Mehdizadeh, A., 2017. Biostratigraphy and corresponding analysis of Middle Jurassic plant
macrofossils of Tabas and Yazd areas. PhD Thesis. University of Ferdowsi, Mashhad (in Persian).
Mehdizadeh, A., Vaez-Javadi, F., Ashouri, A., Ghaderi, A., 2017. Biostratigraphy of Hojedk Formation
in Calshur area, southwest Tabas based on plant macrofossils and palaeoclimate analysis. Journal of
376 Vaez-Javadi et al.
Kharazmi Geosciences, 4 (2): 213-240.
Mehdizadeh, A., Vaez-Javadi, F., Ashouri, A., Ghaderi, A., 2018. Biostratigraphy of Hojedk Formation
in Chahrekhneh, southwest Tabas based on plant macrofossils and palaeoclimate analysis.
Geoscience Researches, 43: 89-112.
Naugolnykh, S.V., 2009. A new fertile Neocalamites from the Upper Permian of Russia and
equisetophyte evolution. Geobios, 42: 513-523.
Navidi Izad, N., Sajjadi, F., Dehbozorgi, E., Hashemi Yazdi, F., 2014. Palynostratigraphy and
sedimentary palaeoenvironment of Dalichei Formation in the Dictash stratigraphy section, north east
Semnan. Journal of stratigraphy and sedimentology researches, 57 (4): 21-46 (in Persian).
Osborn, J.M., Taylor, T.N., 1993. Pollen morphology and ultrastructure of the Corystospermalespermineralized
in-situ grains from the Triassic of Antarctica. Review of Palaeobotany and
Palynology, 79: 205-219.
Phillip, J., Floquet, M., 2000. Late Cenomanian (94.7-93.5). In: Dercourt, J., Gaetani, M., Vrielynck,
B., Barrier, E., Bijuduval, B., Brunet, M.F., Sandulescu, M. (Eds.), Atlas Peri-Tethys Palaeo-
Geographical Maps. CCGM/CGMW, Paris, pp. 129-136.
Phipps, D., Playford, G., 1984. Laboratory techniques for extraction of palynomorphs from sediments.
Department of Geology, University of Queensland, 11: 1-29.
Poole, I., Mirzaie-Ataabadi, M., 2005. Conifer Woods of the Middle Jurassic Hojedk Formation
(Kerman Basin) Central Iran. Iawa Journal, 26 (4): 489-505.
Potonié, R., 1958. Synopsis der Gattungen der Sporae dispersae II. Teil: Sporites (Nachtraege), Saccites,
Aletes, Preacolpates, Polyplicates, Monocolpates. Beihefte zum Geologischen Jahrbuch, 31: 114 pp.
Rees, P.M., 1993. Caytoniales in Early Jurassic Floras from Antarctica. Geobios, 26: 33-42.
Reiser, R.F., Williams, A.J., 1969. Palynology of the Lower Jurassic sediments of the northern Surat
Basin, Queensland. Geological Survey of Queensland, Publication, 339, Palaeontological Paper, 15.
Rossetti, F., Nasrabady, M., Vignaroli, G., Theye, T., Gerdes, A., Razavi, M.H., Moin Vaziri, H., 2010.
Early Cretaceous migmatitic mafic granulites from the Sabzevar range (NE Iran): implications for
the closure of the Mesozoic peri-Tethyan oceans in central Iran. Terra Nova, 22 (1): 26-34.
Sajjadi, F., Hashemi, H., Dehbozorgi, A., 2007. Middle Jurassic palynomorphs of the Kashafrud
Formation, Koppeh Dagh Basin, northeastern Iran. Micropaleontology, 53: 391-408.
Sajjadi, F., Dermaneki Farahani, S., 2007. Palynostratigraphy and palaeoecology of the Middle Jurassic
sediments in southwest Maragheh based on terrestrial palynomorphs. Journal of stratigraphy and
sedimentology researches, 67 (3): 41-64 (in Persian).
Scanu, G.G., Kustatscher, E., Pittau, P., 2015. The Jurassic flora of Sardinia-A new piece in the
palaeobiogeographic puzzle of the Middle Jurassic. Review of Palaeobotany and Palynology, 218:
80-105.
Şengör, A.M.C., 1979. Mid-Mesozoic closure of Permo-Triassic Tethys and its implications. Nature,
279 (5714): 590-593.
Şengör, A.M.C., 1990. A new model for the late Paleozoic-Mesozoic tectonic evolution of Iran and
implications for Oman. Geological Society of London, Special Publications, 49 (1): 797-831.
Shahabpour, J., 1998. Liesegang blocks from sandstone beds of the Hojedk Formation, Kerman, Iran.
Geomorphology, 22 (1): 93-106.
Srivastava, S.K., 1987. Jurassic spore-pollen assemblages from Normandy (France) and
Germany. Geobios, 20: 5-79.
Steart, D.C., Spencer, A.R.T., Garwood, R.J., Hilton, J., Munt, M.C., Needham, J., Kenrick, P.,
2014. X-ray Synchrotron Microtomography of a silicified Jurassic Cheirolepidiaceae (Conifer) cone:
histology and morphology of Pararaucaria collinsonae sp. nov. PeerJ, 2: 1-29. DOI:
10.7717/peerj.624.
Takin, M., 1972. Iranian geology and continental drift in the Middle East. Nature, 235 (5334): 147-150.
Taylor, E.L., Taylor, T.N., 2009. Seed ferns from the Late Paleozoic and Mesozoic: any angiosperm
ancestors lurking there? American Journal of Botany, 96: 237-251.
Taylor, T.N., Cichan, M.A., Baldoni, A.M., 1984. The ultrastructure of Mesozoic pollen -Pteruchus
dubius (Thomas) Townrow. Review of Palaeobotany and Palynology, 41: 319-327.
Taylor, E.L., Taylor, T.N., Kerp, H., Hermsen, E.I., 2006. Mesozoic seed ferns: old paradigms, new
discoveries. Journal of the Torrey Botanical Society, 133: 62-82.
Taylor, E.L., Taylor, T.N., Krings, M., 2009. Paleobotany: The Biology and Evolution of Fossil
Geopersia 2024, 14(2): 361-377 377
Plants. Academic Press, 2nd edition, 1252 pp.
Thomas, H.H., Seward, A.C., 1925. Chapter VI. The Caytoniales, a new group of angiospermous plants
from the Jurassic Rocks of Yorkshire. Philosophical Transactions of Royal Society of London, Series
B, 213: 299-363.
Tipper, G.H., 1921. The geology and mineral resources of eastern Persia. Records of the Geological
Survey of India, 53: 51-80.
Vaez-Javadi, F., 2014. Triassic and Jurassic Floras and Climate of Central-East Iran. Geological Survey
of Iran, Rahi publication, Tehran, 254 pp.
Vaez-Javadi, F., 2015. Plant macrofossils and Biostratigraphy of the Calshaneh section, NW Tabas and
its palaeoclimate analysis. Journal of Stratigraphy and Sedimentology Research, 61 (4): 105-123 (in
Persian).
Vaez-Javadi, F., 2017a. Palynostratigraphy of the Middle Jurassic sediments in Hojedk Formation,
Tabas Block, East-Central Iran. The Palaeobotanist, 66: 60-47.
Vaez-Javadi F., 2017b. Palynomorphs and Plant macrofossils biostratigraphy of the Calshaneh area,
NW Tabas: Palaeoclimate and paleogeography analysis. First International Congress on Jurassic and
Neighboring Countries, Mashhad.
Vaez-Javadi, F., 2018. Middle Jurassic flora from the Hojedk Formation of Tabas, Central East Iran:
Biostratigraphy and palaeoclimate implications. Rivista Italiana di Paleontologia e Stratigrafia, 124
(2): 299-316.
Vaez-Javadi, F. 2020. Middle Jurassic palynology of the southwest Tabas Block, Central-East Iran.
Palynology, 44 (3): 551-562.
Vaez-Javadi, F., Mirzaie-Ataabadi, M., 2006. Jurassic plant macrofossils from the Hojedk Formation,
Kerman area, east-central Iran. Alcheringa, 30: 63-96.
Vaez-Javadi, F., Namjoo, S., 2016. Plant Macrofossils and Biostratigraphy of the Hojedk Formation at
the North Kouchekali, West Tabas and its Climate analysis. Paleontology, 3 (2): 220-243 (in
Persian).
Vaez-Javadi, F., Abbasi, N., 2018. Middle Jurassic biostratigraphy of plant macro and microfossils in
the South Zanjan, Soltanieh Mountains, NW Iran. Geosciences, 106: 91-102.
Vaez-Javadi, F., Mirzaie-Ataabadi, M., 2019. Middle Jurassic plant macro and microfossils from
Shahreza, South West Isfahan, Central Iran: Palaeoclimate influences. Geopersia, 9 (1): 169-193.
Vaez-Javadi, F., Ghanbarian, M.A., 2021. Jurassic palynostratigraphy of the Karim-Abad section, North
of Bahabad, Yazd Province: relative abundance and SEGs analysis. Geosciences, 118: 3-14.
Vaez-Javadi, F., Abbaszadeh, M., 2022. Biostratigraphy of plant macrofossils of the Mazino, southwest
Tabas, Central East of Iran and its palaeoecological analysis. Geosciences, 126 (4): 209-220 (in
Persian).
Vakhrameev, V.A., 1991. Jurassic and Cretaceous floras and climates of the Earth. Cambridge
University Press, England, 318 pp.
Van Konijnenburg-van Cittert, J.H.A., 1971. In situ gymnosperm pollen from Middle Jurassic of
Yorkshire. Acta Botanica Netherland, 20 (1): 1-97.
Van Konijnenburg-van Cittert, J.H.A., van der Burgh, J., 1989. The flora from the Kimmeridgian (Upper
Jurassic) of Culgower, Sutherland, Scotland. Review of Palaeobotany and Palynology, 61: 1-51.
Van Konijnenburg-van Cittert, J.H.A., van der Burgh, J., 1996. Review of the Kimmeridgian flora of
Sutherland, Scotland, with reference to the ecology and in situ pollen and spores. Proceedings of the
Geologists' Association, 107 (2): 97-105.
Van Konijnenburg-van Cittert, J.H.A., 2002. Ecology of some Late Triassic to Early Cretaceous ferns
in Eurasia. Review of Palaeobotany and Palynology, 119:113-124.
Van Konijnenburg-van Cittert, J.H.A., Kustatscher, E., Pott, C., Dutsch, G., Schmeissner, S.,
2017. First record of the pollen-bearing reproductive organ Hydropterangium from the Rhaetian of
Germany (Wustenwelsberg, Upper Franconia). Neues Jahrbuch für Geologie und Paläontologie, 284:
139-151.
Vassiliev, Y., 1984. Mesozoic plant fossils from coal areas in Iran. 2: 97 pp. (translated into Persian by
Mehdian, M.H.). Atlas of the Ministry of Mine and Metal, 2(2): 47 pls. Tehran.
Vijaya, K., Sen, K., 2005. Palynological study of the Dubrajpur formation in the Mesozoic succession,
Pachambi Area, Birbhum Coalfield, West Bengal. Journal of the Palaeontological Society of India,
50 (1): 121-133.
378 Vaez-Javadi et al.
Watson, J., 1988. The Cheirolepidiaceae. In: C.B. Beck (ed.), Origin and Evolution of Gymnosperms.
pp. 382 - 447. Columbia University Press, New York.
Weiss, M., 1989. Die Sporenfloren aus Rät und Jura Südwestdeutschlands und ihre Beziehungen zur
Ammoniten-Stratigraphie. Palaeontographica B, 215 (1-6): 1-168.
Zanchi, A., Zanchetta, S., Berra, F., Mattei, M., Garzanti, E., Molyneux, S., Namab, A., Sabouri, J.,
2009. The Eo-Cimmerian (Late? Triassic) orogeny in North Iran. Geological Society of London,
Special Publications, 312 (1): 31-55.
Zanchi, A., Malaspina, N., Zanchetta, S., Berra, F., Benciolini, L., Bergomi, M., Cavallo, A., Javadi,
H.R., Kouhpeyma, M., 2015. The Cimmerian accretionary wedge of Anarak, Central Iran. Journal
of Asian Earth Sciences, 102: 45-72.
Zavialova, N., van Konijnenburg-van Cittert, J.H.A., 2016. Exine ultrastructure of in situ pollen from
the cycadalean cone Androstrobus manis Harris, 1941 from the Jurassic of England. Review of
Palaeobotany and Palynology, 225: 33-42.
Zhang, J., Lenz, O.K., Wang, P., Hornung, J., 2021. The Eco-Plant model and its implication on
Mesozoic dispersed sporomorphs for Bryophytes, Pteridophytes, and Gymnosperms. Review of
Palaeobotany and Palynology, 293: 1-28.
)